The City in the Shadow of Prism Tower in Pokemon Legends Z-A is the fifth main mission, which introduces you to some in-game mechanics and allows access to some Wild Zones featuring new Pokemon. It is one of the longer missions in-game as it includes multiple cutscenes along with a few side quests. Moreover, you receive an invaluable item while completing it.
Here's a complete walkthrough for The City in the Shadow of Prism Tower mission in Pokemon Legends Z-A to help you clear it easily.
How to complete The City in the Shadow of Prism Tower mission in Pokemon Legends Z-A
Begin the mission at Hotel Z

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The City in the Shadow of Prism Tower mission in Pokemon Legends Z-A automatically begins from where the fourth mission ends after your victory against Zach. Once you are finished celebrating the win with other Team MZ members, leave Hotel Z.
As you leave, a cutscene will trigger, beginning the fifth main mission. During the cutscene, you will receive an important item called Red Canari Plush from Tarragon, which can be equipped to your critter to increase the experience they earn. Moreover, you will be informed about a collectible item called Colorful Screws.
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Exploring Lumiose City
Afterward, follow your rival (Taunie/Urbain) from Hotel Z on a linear route. Along the way, Mega Crystals will appear. You can break these with your Pokemon to collect Mega Shards, which can be traded for Mega Stones (useful to trigger Mega Evolutions) and Exp Candies later in the game.

During this part of the quest, you will also learn about the Rotom Phone’s safety mode, which allows you to jump from heights freely. You can use this ability to keep following your rival towards Nouveau Cafe. Along the way, you can explore Wild Zone 2 to catch local Pokemon. This is a good place to add critters to your party.
Following this, you will encounter Vinnie, the secretary to Quasartico’s CEO.
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Fist of Justice challenge
As you continue with The City in the Shadow of Prism Tower mission in Pokemon Legends Z-A, you will eventually reach Wild Zone 3. However, its entrance will be blocked as Ivor, the leader of the Fist of Justice organization, is delivering a speech at the spot.

Meanwhile, Josee from the organization will challenge you to a battle. You must defeat her to progress through this quest.
Josee's team comprises a Machop and a Meditite. Since both of them are Fighting-type Pokemon that are weak to Flying-types, it is recommended you use a monster like Pidgey or Fletchling that can use Flying-type moves to defeat them.
Following your victory, a cutscene will play out where the Fist of Justice members will leave, and you can access Wild Zone 3.
Note that the next part of this main mission requires you to have a Pikachu. As such, you must catch one at Wild Zone 3. Additionally, if you wish Pikachu to be one of your party members, obtain two of them.
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The Looker Bureau and side missions

After catching Pikachu, head over to the Looker Bureau for a cutscene to play out. Following that, talk to Emma to undertake three side missions. You must complete each one to proceed forward in The City in the Shadow of Prism Tower mission. Here's how you can clear each of the side missions:
Side Mission 001: A Big Ol’ Bunnelby
- Go to the alley with the client and defeat an Alpha Bunnelby. You can also catch it.
- Rewards: 400 Poke Dollars, 2x Fresh Water
Side Mission 002: A Use for an Evolution Stone
- Trade a Pikachu for a Heracross with the client.
- Rewards: 800 Poke Dollars, 5x Poke Balls, 3x Potion.
Side Mission 003: Sableye in the Cemetery
- Enter Wild Zone 4, navigate to the top-right, defeat a Sableye, and retrieve the client’s ring.
- Rewards: 4000 Poke Dollars, 2x Revive.
Once all three tasks are complete, return to Emma and interact with her to trigger a cutscene where she Mega Evolves her Ampharos.
Once that is finished, The City in the Shadow of Prism Tower mission will be completed in Pokemon Legends Z-A, and the next mission, Reaching Rank X, will begin.
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